Pine cones are eaten by both red and grey squirrels. Pine cones are also consumed by some bird species, such as the crossbill and woodpecker. Pine cones are consumed by squirrels, who then discard the cores and removed scales.
Some birds also leave piles of stripped pine cone scales behind, however unlike the scales that squirrels leave behind, which often have neat edges, the scales that birds leave behind are typically rough from being torn from the cone.
To make a pine cone simpler to chew, woodpeckers will occasionally press the cone against the bark of a tree. The consistent accumulations of pine cone waste left beneath trees serve as proof.